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Plans for the Yellowstone grizzly

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Grizzly bear on Swan Lake Flats, Yellowstone

Grizzly bear on Swan Lake Flats, Yellowstone

Grizzly bears have been in the news in recent years. First, because human-bear conflicts have been more numerous, including a total of six deaths of people since 2010. Managing these conflicts and the bear predation on cattle means about twenty grizzlies are intentionally killed or removed to zoos per year (see this database if you are interested). In 2016 the news is that the U.S. Fish and Wildlife Service is proposing to take them off the Endangered Species list by the end of the year. The National Park Service put together an informative page about the history of efforts to help the Yellowstone-area grizzly recover, including listing it and delisting it, plus explaining how to minimize encounters with bears and ensure your safety if you do accidentally come close to one. You can find the article here: “Grizzly Bears and the Endangered Species Act.” The most recent delisting was in 2007, but a court ruling overturned that and put them back on the threatened species list in 2009.

Here’s a quick list of safety points, courtesy of the NPS. When backcountry hiking, you can reduce the odds of being injured by a bear by following these five rules:

  1. Hike in groups of three or more people.
  2. Stay alert.
  3. Make noise in areas with poor visibility.
  4. Carry bear spray.
  5. Don’t run during encounters with bears.

The grizzly bear population has made a remarkable recovery, to about 700 individuals in the Greater Yellowstone Ecosystem. So why is delisting controversial? Some are worried about plans for hunting in the surrounding states of Idaho, Montana, and Wyoming. For a March 20, 2016, article that quotes the state governors on the subject, see “US seeks end to Yellowstone grizzly protections” on the Explore Big Sky website.

Do you have an opinion on this subject? Isn’t this photo beautiful? Write your informed comment below.
—Editor Beth

Photo by Jim Peaco for the National Park Service, June 2005.

What is meant by “Greater Yellowstone”?

Categories: Flora and Fauna, Park environs, Wildlife
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Back in 1872, when Congress was wise enough to set aside a large area of “useless” land and name it the Yellowstone National Park, the main purpose was to reserve the remarkable geothermal features and their surroundings “for the benefit and enjoyment of the people.” This phrase was later inscribed on the park’s North Entrance Arch. Park boundaries were set to correspond to obvious natural features and partially along lines of longitude and latitude.

Protecting wild animals was not a big concern in 1872, since there seemed to be a great many of them. But it soon became apparent that they needed protection from overzealous hunters. Concern for the buffalo (now usually called bison) was great. They were rapidly being wiped out, largely because of government policy that rewarded buffalo hunters richly. Native American Plains Indians depended upon the buffalo for their very livelihood, but U.S. officials wanted the tribes to settle on reservations and make life safer for Eastern homesteaders.

Bison herd in winter, by Bruno Giletti, "Yellowstone Treasures," page 343

Bison herd in winter, by Bruno Giletti, “Yellowstone Treasures,” page 343

Even before bison were nearly driven to extinction, people who cared about preserving wildlife and the western lands from rampant development moved to set aside more land for special protection. In 1891 the Department of the Interior created the Yellowstone Park Forest Reservation (now part of Shoshone National Forest), paving the way for today’s 155 national forests.

Now we skip ahead one century plus a year after Yellowstone Park was designated. In 1973 concerned thinkers and planners convinced Congress to pass the Endangered Species Act. By then, bison were thriving but the grizzly bears were not;

Grizzly bear, Courtesy of NPS, "Yellowstone Treasures," page 344

Grizzly bear, Courtesy of NPS, “Yellowstone Treasures,” page 344

local agencies also found by the 1970s that cutthroat trout, pronghorn (also called antelope), whitebark pine, and quaking aspen trees were of special concern. An area of about four million acres with Yellowstone at its center was christened the Greater Yellowstone Ecosystem. In 1986 the federal government recognized Greater Yellowstone, when a joint congressional committee report outlined shortcomings in interagency coordination and concluded that the area’s essential values were at risk.

Exact boundaries of the GYE are hard to define, and they  keep changing over the years.

Exact boundaries of the GYE are hard to define, and they keep changing over the years. Map from Greater Yellowstone Coalition website.


The National Park System now describes the GYE thus:
• 12–22 million acres; 18,750–34,375 square miles (Sizes, boundaries, and descriptions of any ecosystem can vary.)
• States: Wyoming, Montana, Idaho
• Encompasses state lands, two national parks, portions of five national forests, three national wildlife refuges, Bureau of Land Management holdings, private and tribal lands
• Managed by state governments, federal government, tribal governments, and private individuals.

In addition to government agencies like the Interior Department’s National Park Service and the Forest Service (part of the Department of Agriculture), a number of nonprofit agencies work to help preserve Greater Yellowstone; National Wildlife Federation, Defenders of Wildlife, and the Greater Yellowstone Coalition are three of them. Two prime concerns of all these entities are climate change and providing corridors for wildlife migrations.

Another spectacular mountain area farther north is the Crown of the Continent, an initiative spearheaded by University of Montana geography professor Rick Graetz with the assistance of his wife Susie Graetz and others. Covering the area centered upon Waterton-Glacier International Peace Park, it extends some 250 miles from Alberta, Canada south along the Continental Divide into central Wyoming.

A stated goal of this coalition is to maintain “wildlife corridors [that] may mean the difference between a robust grizzly population and one needing continued human protections, particularly in an age when movement will be essential for both grizzlies and other species that might need to head north to weather the impending climate storm.”

Educators and governmental agencies in this entire area are stressing the interconnectedness of the ecology of this beautiful mountainous area all the way from Alberta, Canada to west-central Wyoming. I was interested to find that western North America is not the only part of the world pondering this question. An Australian website states: “Wildlife corridors can range in size – from small corridors created by local communities to large corridors that stretch across many different landscapes.

“For example, a small corridor might be an area along a creek that has been revegetated by a local community group to link two patches of forest. Native animals could then move more freely between these forests to find food, shelter and opportunities to breed.

“Large-scale corridors might span tens or hundreds of kilometres across multiple landscape types and jurisdictions. Typically a large-scale corridor would require collaboration between a wide range of groups working in partnership to manage them.”

A puzzle I have not been able to solve in my research into the two U.S. entities is whether they would wish to connect the Crown of the Continent with the Greater Yellowstone Ecosystem, since there’s a large part of Montana between them where many people live. Grizzlies and wolves are not compatible with what we consider civilization. Perhaps just making more people aware of the beauties of the two areas, the threats to the species we share them with, and how we can help preserve them is enough for now.

Observing the American dipper

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(Part II of Billy Hofer’s article, continued from yesterday)

[The American dipper’s] plumage is dense and compact, and is moreover well protected by a coat of oil, which he often renews from the large oil sack with which he is provided. So the dipper never gets wet, and we may conclude never takes cold, although he passes the winter about the open places of mountain torrents as far north as Alaska. His food, which he secures from the bottom of the stream, consists mainly of aquatic insects, and although his first cousin who inhabits Northern Europe has been accused of feeding on the spawn of trout and salmon, there is not a particle of evidence to sustain the grave charge.

The dipper dives into the water and gets to the bottom as soon as possible, and by means of rapid wing beats and holding on to the stones and gravel with his feet resists the constant tendency to shoot up through the water to the air above. He vigorously turns over the stones and sticks, and secures the water insects and aquatic larvae which are found in such situations, and then when he requires breath comes to the surface again. He presents a queer appearance when at the bottom, where he seems to be tumbling about in the most curious and erratic way, now for a moment stationary, and again being swept away by the force of the stream. No doubt he makes a good living, for he always seems busy, contented and cheerful.

The dipper seems to be satisfied with his own company for the greater part of the year, but during the breeding season he perforce seeks a mate, and these two build their nest close to the water’s edge, under a bank or the roots of a tree, and often amid the spray of some brawling cascade. The structure is as curious in its way as its architects. It is lined with twigs or fine, dry grass, and completely covered on the outside by an arched roof of fine green moss, which is kept fresh and living by the moisture of its surroundings, or sometimes perhaps by the birds themselves, who, it is said, after emerging from the water, fly to the roof of the nest and, alighting there, vigorously shake the drops from their feathers over the mossy covering.

– – – – –
I am quoting this from Hofer’s “Through Two-Ocean Pass,” Part XII, April 9, pages 202–3, Forest and Stream magazine, in fifteen parts, January 29 through April 30, 1885.

A delightful bird found in Yellowstone, the American dipper

Categories: Flora and Fauna, Through Early Yellowstone, Wildlife
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(Part I)
There’s a rather small and plain bird that I’ve loved to watch whenever I’m lucky enough to be in the Western mountains. Formerly called the water ouzel, the American dipper (Cinclus mexicanus) shows up on pages 337–38 of Yellowstone Treasures in a quote from Owen Wister. But there’s a longer story I came across in an old Forest and Stream magazine.
320px-American_DipperPhoto from Wikipedia Commons

The winter adventurer, Billy Hofer—whose story of an incredible 1887 ski trip through Yellowstone will appear in my forthcoming anthology, Through Early Yellowstone—must have done some research on dippers, although he does not tell us that. His delightful story about them, written two years before his winter trip, is one that all bird watchers can appreciate. Here’s what he wrote 130 years ago.

[American dippers are] curious little drab-colored birds, progressing with an odd bobbing motion along the water’s edge, or taking short flights from one almost submerged rock to the next.

[The bird ranges from] Alaska to Mexico, and it only insists on clear streams among the mountains. On such brooks and rivers as fulfill these conditions the dipper is found in abundance—though it is not a particularly sociable bird, and seems rather to prefer to be alone—and its cheerful song and remarkable habits render it a pleasing feature alike of the dark cañons and of the open sunlit glens.

One feels a ludicrous sensation of astonishment when first observing this species. It is not very surprising to see a bird sitting upon the water or flying above it, dive beneath its surface, but it is really startling to see one calmly walk down a shelving rock or a smooth beach into the water, and keep on without any apparent attempt at swimming or diving, until it has disappeared. It wholly upsets one’s ideas of specific gravity, for we are accustomed to think that birds and mammals will naturally float, and that to dive or disappear beneath the water requires some apparent effort. It is not so with the dipper, however. He marches deliberately into the turbulent water, which engulfs him and usually sweeps him several feet down the stream before he bobs up serenely to the surface, and either takes wing or sculls himself ashore. It must not be supposed that he always enters the water in this way. Quite as often he dives from the wing or from the surface of the stream, but perhaps the method which he likes best is to plunge from the top of a high rock or a log a foot or two above the current, and then, after his battle with the torrent, return to the same perch, from which at intervals he trills his simple though very sweet song.

His method of progression under water is like that of most, if not all, diving birds, by means of the wings. His feet are not better adapted fro swimming than those of a robin, and although he gets along capitally when sculling about the edges of the mountain holes, they would be of little service to him under water except sometimes as anchors. He flies under the water therefore with nearly open wings, and gets about very actively.

(To be continued tomorrow.)

Bear safety

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Grizzly bear from page 344 of Yellowstone Treasures, 4th ed.

Grizzly bear from page 344 of Yellowstone Treasures, 4th ed.

In a recent press release about preparing for fall, Yellowstone National Park rangers remind us that the park is bear country. Here’s their advice.

In the fall, grizzly bears and black bears usually move to higher elevations to feed on whitebark pine seeds, and consume the calories they need to sustain themselves during winter hibernation, but they may be encountered along roads or hiking trails throughout the park. When hiking or backpacking, remember to travel in groups of three or more, make noise on the trail, and be alert for bears. All hikers should always carry bear spray so that it is readily accessible—not inside a pack—and know how to use it. Bear spray is proven to be highly successful at stopping aggressive behavior in bears. It is sold at bookstores, gift shops, outdoor stores, and service stations inside the park, as well as in many stores in the surrounding communities. New this year, bear spray is now available for rent at Canyon Village in a kiosk near the Canyon Visitor Education Center through late September.

Park regulations require people to stay a minimum of 100 yards (the length of a football field) away from bears and wolves at all times. If you see a bear along the road, move off the road and park on the shoulder or in a pullout and stay in your vehicle to watch the bear. Use your binoculars, telescope, or telephoto lens to get a closer look at the bear rather than approaching the bear.

Happy wildlife watching, and stay safe!
—Beth Chapple, editor at Granite Peak Publications

Photo credit: Department of the Interior, National Park Service, Yellowstone National Park

What are the different species of mammals in Yellowstone Park?

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On the website Quora.com, someone recently asked about the mammals one sees in Yellowstone Park, so I tried my hand at an answer. I’m not a real animal person, being more enamored with things that stay in one place like geysers, hot springs, flowers, lakes, waterfalls, and mountains, yet I have personally seen all but the rarer animals in my many visits to Yellowstone.

The big mammals—sometimes called “charismatic megafauna” with tongue in cheek—are the bison (around 4000 currently), the elk (8,000 to 10,000), the moose (very scarce, and more often seen in neighboring Grand Teton National Park), and both grizzly and black bear (several hundred of each).

The gray wolf population is hovering around 100. Mule deer are much more common than white-tailed deer. Then there’s the pronghorn (commonly but incorrectly called antelope)—my favorite animal for being so beautiful, graceful, and fast. Their population fluctuates around a couple of hundred seen in northern Yellowstone, as are bighorn sheep.

Pronghorn

Pronghorn

Coyotes are very common, less so the red foxes and mountain goats (the latter considered to be migrants to the park). Mountain lions, bobcats, and especially lynx are there but almost never seen.

Small carnivores you might see include badgers, river otters, and raccoons. Then there are the most common rodents: yellow-bellied marmots, Uinta ground squirrels, muskrats, red squirrels, and the tiny pikas or coneys. Beaver have become more numerous in recent years.

Pika

Pika

And this is just a sampling. There are many smaller mammals like voles, mice, bats, and shrews.

What’s New, Fun, and Interesting in Yellowstone This Summer?

Categories: Geysers, Trip Reports, Wildlife
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Entering Yellowstone from the North Entrance may be a little tough going and not aesthetically pleasing for most of this year [2015], since there’s a humongous construction project going on to completely revamp the entrance area at the little town of Gardiner. But five miles and a thousand feet up the road to the south is Mammoth Hot Springs, and, in addition to seeing the springs along the Upper Terrace Road, I recommend spending an hour or so at the redone Albright Visitor Center. It has excellent hands-on dioramas of all of the park’s bigger mammals and kiosks for park orientation on the first floor. In the basement level, completely accessible with a new elevator, are great historical displays and the restrooms. For more about this see the Yellowstone Insider’s recent article.

One of Upper Geyser Basin’s most popular sites is the wonderfully regular Riverside Geyser. It almost always erupts every six to six-and-one-half hours. Here is the eruption I caught on my all-too-short visit to the park in mid June.


You can hear (1) a geyser gazer transmit by FRS radio the time of eruption to the Old Faithful Visitor Center, (2) the excited crowd,(3) the swishing of the main eruption, and (4) the rumbling of the side spouter that always accompanies Riverside’s eruptions. It always erupts quite a bit longer than this little video, which was edited for Granite Peak Publications by Jens Paape.

You can reach Artemisia Geyser’s beautiful pool and formation in one of two ways.Artemisia Geyser One is by walking beyond Riverside Geyser about half a mile up what used to be the main road and is now a rather rough trail past Morning Glory Pool (page 95 in Yellowstone Treasures) or by parking at Biscuit Basin and crossing the road to reach the other end of the trail from Morning Glory Pool. Up the hill in the distance in my picture is Hillside Springs, which old-time stagecoach drivers called Tomato Soup Springs.

I did not see any grizzly bears on this trip, but there are now enough of them in the Greater Yellowstone Ecosystem so that visitors are seeing them quite frequently. The national media covered the recent very unusual event where a grizzly climbed on the hood and sides of an occupied car, leaving some scratches but giving the occupants of the car the thrill of their lifetime and their own video.

One thrill of this visit for me was being assigned for three nights to what has to be the best room in the Old House of Old Faithful Inn (Room 229). It was inside the farthest east of the five dormer windows that span the third floor front of the inn. Two mornings I awoke to a swishing sound, opened the side window, and there was Old Faithful Geyser erupting for my private enjoyment!

For fishermen and others interested in what is happening with the fish in Yellowstone Lake these days, take a look at the Great Falls Tribune’s story about the good news regarding the struggle against illegally introduced lake trout.

For wolf devotees and other wildlife lovers

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For a small investment, you can watch from afar the comings and goings of Yellowstone Park’s wildlife. This is a worthwhile website with reports by real naturalists and experienced wolf-watchers.

Besides the almost daily reports, you can also find pictures of the animals and birds the naturalists are seeing. Spring is a great season to be watching all this! And it’s a time when—for personal reasons—very few of us are likely to be there. In fact, these weeks are also not the best time to visit most of the park, because not all roads and facilities are open yet, and because the weather is usually quite iffy until mid June or so.

Guest Post by Rita Reining, a nature lover and outdoor enthusiast. She can often be found hiking the hills around her home in Oakland, CA and serves as a docent in the Natural Sciences Gallery of the Oakland Museum of California.

When my friend Ellen and I decided to sign up for Wolf Week—a five-day course presented by the Yellowstone Association at the Lamar Buffalo Ranch the last week in March—I had no idea of the adventure that was in store.

After dinner on our first evening the instructors gave out a picture and a short bio of a wolf living on the North Range of the park to each pair of participants. Each couple presented their wolf information to the group. We learned about the 11 different packs in the park, but we concentrated on the eight packs that currently call the North Range home. The specific wolves we learned about all have radio collars. The wolves generally don’t have names but are known by their number. Also, we learned a little of the genealogy of the wolves, many of whom are descendants of the Druid pack which were the original wolves introduced to Yellowstone in 1995. We felt a bit better acquainted with specific wolves and the packs that we might encounter.

The next morning (and for the following two mornings) we were up at 5 AM in order to be on the bus to start looking for the wolves before sun-up. We drove to the Little America area where we came upon a group of people with spotting scopes focused on the Junction Butte (JB) pack about one-half mile away up the hillside enjoying the remains of an elk brought down a couple of days earlier. We quickly set up our scopes to observe our first wild wolves of this trip. It was so exciting!

As we watched them, I was surprised that their behavior was quite sociable. When we arrived, the big gray alpha male, 911M, was eating. A bit later the alpha female, 970F, and then two other females settled in to feed. No snarling or fighting such as I had expected. When it seemed that all had had their fill, the pack began to howl. First one, then another, then everyone. Each wolf has its own tone and voice. It was an exhilarating chorus. From across the way, more howling was heard. This was by the Prospect Peak pack (PP). The JBs evidently did not want to interact with the PPs and they ran up the hill and disappeared over the ridge.
Wolf March 2015 by Rita Reining
There are several groups of dedicated wolf watchers. First there are those who are associated with The Wolf Project, consisting of teams who follow all the packs in the park. Then there are the rangers, photographers, bloggers, and local residents who are out every day recording and reporting what they see. All seem to be connected by radio, and we relied on this communication every day in order to follow the wolves.

Thus, we learned that there was wolf activity in the Lamar Valley. This time we watched a gray pup of the Lamar Canyon (LC) pack busy at an old bison carcass. Soon the black alpha female, 926F, walked by, stopped for a moment by the carcass and continued on her way, followed by another black pup.

We followed 926F and her six pups’ activities during our wolf week. The picture above was taken by Jane Morse, a fellow participant, and this wolf looks very much like 926F. The stories about 926F made our observations all the more interesting.

About ten days before we arrived in Yellowstone, the LC alpha male was killed by the PPs. (The highest cause of wolf deaths is by other wolves.) The larger PP pack encountered the smaller LC pack. The reason for the PP aggression is unknown, but as they were advancing on the LCs, all the LCs ran away except the alpha male. He stood his ground as the PPs edged closer, and then he turned and ran away in the opposite direction from the rest of his pack. Unfortunately the PPs caught and attacked him, leaving him for dead. 926F came back and lay by him as he died.

Now 926F had become the leader of the LC pack, consisting only of a pregnant 926F and her six yearling pups. She was solely responsible for getting food for herself and her pups. Wolves only begin to learn to hunt after their first year. 926F will need a new mate. All the wolf watchers were concerned for the future of the LC pack. 926F would be denning soon to have her new pups. Mother wolves do not leave the den until the pups can be left without her for a short time. Usually the alpha male stays near her, and he and other members of the pack bring her food. The LC wolves were in danger of starving, since the yearling pups were useless as hunters.

The next morning we saw a deer carcass in the ditch by the road. It had been hit by a car the night before. Here was a free meal, and 926F walked right by a small herd of bighorn sheep to get to the carcass. But she ran off when the rangers came to move the carcass away from the road for the safety of the wolves as well as to avoid a traffic jam. As soon as the rangers left, a golden eagle, some magpies, and a coyote returned to the carcass. But, 926F wasn’t going to let the lesser beings take food away from her and her pups, so about noon she came and dragged the carcass into the woods.

On the final morning in the field we went back to where we had last seen 926F. We had only just left the bus when we heard her howling. Shortly, her pups joined in. The howling had the bighorn sheep across the road on high alert, but we couldn’t spot any of the wolves. About a half hour later, we saw them crossing the ridge under a cliff. A lone bull elk was standing at attention on an outcropping as they passed by him within 100 yards. The pack seemed to ignore him. Two elk were on top of the cliff, on alert, scanning in opposite directions. Just then we saw one of the pups approaching the elk. Both elk turned and faced the wolf. The wolf got to about 50 feet from the elk, stopped, backed up a couple of paces and gave the elk a wide berth as he walked around them. Then we saw him walking toward the elk from uphill. He more or less repeated his previous approach from below, then walked away. Maybe he was just practicing for future hunts. This was the closest we came to seeing any real drama on this trip.

The pack was moving west, so we went west to the other side of the ridge. We spotted the pack bedded down at the edge of the woods. After a while they began to move around. I watched as one of the pups laid on his back in the snow rolling back and forth, just like my dog does. Making wolf “angels”?

Then 926F must have given some signal. She started moving west again at a determined pace with the pups following. We watched until they were out of sight.

All too soon our week at the Lamar Buffalo Ranch ended. We left Yellowstone hoping to sign up in the future for another exciting and interesting course with the Yellowstone Association. Also, I was wondering what the future would be for 926F.

Post script for 926F: Shortly after my visit, four adult males from another pack joined the LCs. The wolf watchers were optimistic that 926F would choose her alpha male from one of them and, with the yearling pups, have a sustainable pack. It was not to be. As of a month after I left Yellowstone, I learned that 926F was now alone. Her yearlings were gone and so were all four of the males. The last I heard, 926F had not yet gone to den. She is very lean but has managed so far to survive on her own. Her saga continues.

“The Wild Lab” of Yellowstone Park

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Today I’m thinking about Yellowstone’s animals, the “charismatic megafauna”—an expression that makes me smile. And I’m anticipating being able to present my first-ever guest blog, by my friend Rita Reining. Rita took part in a late season wolf-watching seminar this winter given by the Yellowstone Institute. Their next winter’s classes are not yet listed but will be up by August or September.

YT174 copy
Wolf picture from “Yellowstone Treasures,” page 351

Awaiting her article, I’ll pass on some thoughts inspired by a Science Magazine article, “Lessons from the Wild Lab,” March 20, 2015, pages 1302–7.

A video circulating on the Web a few months ago attributed recent changes in Yellowstone’s environment to the reintroduction of gray wolves, beginning in 1995. The video seemed to claim that the observed changes, all the way through the animal and plant kingdoms to the re-channeling of rivers, were due to predation on elk by wolves. However, according to writer Virginia Morell from Yellowstone, scientists disagree as to whether or not the coming of the wolves was the only trigger for observed changes “in species from elk to coyotes to willows to bison to beavers.” She cites cougars, grizzly bears, and drought as partners in the park’s transformation.

Wolves and cougars were both extirpated by the early 1930s. Coyotes, however, have been protected from 1936 on—a discrimination in the ruling that I have never understood, since these smaller canids can also prey on animals as large as deer or (very occasionally) elk and have even been known to attack humans.

In addition to the larger predators, Douglas W. Smith, the Yellowstone Wolf Project leader, who has studied wolves for more than three decades, says that smaller predators and humans should be factored in to the great depletion of the elk herd since wolves came to the park.

Here are a few megafauna statistics from Morell’s article:

Elk:
1994, more than 19,000 in the northern range of Yellowstone
2008, just over 6,000

Cougars (known in the park):
1972, almost none
1995, estimated at 15 to 20
2015, estimated at about 35, a “natural experiment,” according to Morell, “but they are
rarely seen and do not howl”

Bison:
1997 (after slaughter of 1,000), about 1,600
2004, over 4,000
2014, 4,900

Beaver colonies:
2014, 12 (but the
park website
) gives 112 beaver colonies in 2011.)*

Wolves:
1994, none
2013, 95 in the park; 34 on the northern range

Human visitors:
2014: 3.6 million.

– – – – –
*From the NPS Yellowstone website: “The increase [in beavers] has occurred throughout the park and is likely related to the resurgence in willow since the late 1990s, at least on the northern range, and possibly in the park interior.”